| Education |
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High school leaving examination, 1973,
Aurajoen yhteiskoulu, Turku
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MSc, 1976, Zoology, University of Turku
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Lic Phil, 1976, Zoology, University of Turku
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PhD, 1982, Zoology, University of Turku
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Docent, 1990, Theoretical ecology, University of Lund
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| Main positions |
University of Turku
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Assistant in zoology, 1977-1985, 1986-1991
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Researcher, Academy of Finland 1979-1985
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Lecturer in biology, 1985
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Assistant professor in zoology, 1986
University of Lund
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Assistant in theoretical ecology, 1986-1990
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Associate professor in theoretical ecology 1991-1997
University of Oulu
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Professor in botany, 1996-
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